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A review of sulfur-containing compounds of natural origin with insights into their Pharmacological and toxicological impacts | Discover Chemistry | Springer Nature Link

Particularly, lysyl oxidase, a copper-dependent amine oxidase, is a key enzyme for collagen and elastin cross-linking, which further enhances the cartilage formation (106)

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(2) Catalase was found in membrane and extracellular compartment but the exact nature and function of these catalases are unknown